Pakistan says its intelligence agents have been accused by the US of alerting al-Qaeda linked militants before the US launches missile attacks against them.
Defence Minister Ahmed Mukhtar said members of Inter-Services Intelligence were accused of "tipping off" militants before strikes in the tribal areas.
Mr Mukhtar said that the Americans "mistrusted" the ISI.
His unusual public admission of the rebuke seems to mark a new low in ties between the US and Pakistan's spies.
'Who's in charge?'
Mr Mukhtar was speaking in Washington, where he is accompanying Prime Minister Yousuf Raza Gilani on his first visit to the US.
Speaking on Pakistan's Geo TV, he said the Americans had alleged that information about targets was being "leaked".
"The burning issue of course is as to who controls the ISI," he said.
The ISI is accused of supporting Taleban fighters in Afghanistan
"In their [the Americans'] view there are some people at some level in the ISI who tip off the Taleban at some level about impending missile attacks when these are shared with the Pakistanis.
"They [the Americans] have expressed displeasure over this."
Mr Mukhtar also openly admitted that President Bush had asked who was really in charge of the ISI during the visit to Washington.
The BBC's security correspondent Rob Watson says that relations between the American Central Intelligence Agency (CIA) and the ISI appear decidedly strained.
Our correspondent says that the stern message apparently being delivered to Islamabad was that Pakistan has to do more to tackle ties between the ISI and Islamic extremists based in the country's tribal areas.
American unhappiness with Pakistan's recently-elected civilian government could also embarrass President Musharraf and the Pakistani army, in addition to raising new concerns about the stability of the nuclear-armed state.
'Great institution'
The US no longer gives Pakistan advance notice when it targets militants in tribal areas, correspondents say.
I've been saying it for a long while now: Pakistan isn't helping out, and is actually playing both sides of this war. That's something that shouldn't be tolerated AT ALL. Aid should be shut down, blockades for trade should be enacted, and other types of punishment.
Every time we go after Al-Qaeda's big fish, we seem to just miss them. Wonder no longer as to why. Personally, if they cost us Bin Laden, or any of his henchmen, I think we ought to find creative ways to get even. :)
Travis
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